| Main Ref. | Whitehead, P.J.P., 1985 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | one clear seasonal peak per year |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Spawns in freshwater in spring or early summer, ascending rivers and streams, the young presumably descending in autumn. |
| Search for more references on reproduction | Scirus |