Africa: ranges from Ethiopia in the north, through East and Central Africa, extending westwards into Angola and reaching the southern-most limits of its distribution in Vungu, KwaZulu-Natal, in South Africa (Ref. 2801, 52193). In the Congo basin known from Lake Mweru, Luapula, upper Lualaba, Lufira (Ref. 41590) and upper Lulua (Ref. 42554).