Classification / Names
Common names | Synonyms | Catalog of Fishes(genus, species) | ITIS | CoL | WoRMS | Cloffa
Environment: milieu / climate zone / depth range / distribution range
Freshwater; brackish; benthopelagic; depth range - m (Ref. ), usually - m (Ref. ). Temperate; 4°C - 20°C (Ref. 2059); 49°N - 40°N, 25°E - 54°E
Eurasia: Widespread all over the Marmara, Black and Caspian seas basins; inhabits mostly fresh- and brackish waters with low salinity (<2%).
Length at first maturity / Size / Weight / Age
Maturity: Lm ?, range 6 - ? cm
Max length : 16.2 cm TL male/unsexed; (Ref. 2058); max. reported age: 5 years (Ref. 59043)
(total): 7 - 8;
soft rays: 12 - 16. This species is distinguished from its congeners entering freshwater in Europe by the following characters: irregular position and shape of diagonal bars on body; first branched ray of second dorsal about as long as penultimate ray; no scales on midline of nape, in front of preoperculum; pelvic-disc fraenum with small rounded lobes and the length is less than 1/6 of width at base; scales in midlateral series 54-62 + 2-3; posterior part of first dorsal without black spot (Ref. 59043).
Occurs in brackish- and fresh-water habitats with low salinity (< 2 ppt); lagoons and lakes; large rivers to small, fast-flowing streams; on sand or mud bottom; mainly in well vegetated or high-complexity habitats. Abundant in backwaters and still channels (Ref. 59043). Longevity is 4-5 years; spawns for the first time at 2 years; spawning season in April to June, occasionally until mid-August; females may repeat spawning during a season; usually spawns for a single season. Males guard eggs until hatching; with adhesive eggs deposited on stones, shells and aquatic plants (Ref. 59043). Feeds on crustaceans (esp. Corophiid amphipods), aquatic insects (mostly chironomid larvae), polychaetes, also small fish and mollusks (Ref. 4696).
Adhesive eggs are deposited on stones, shells and aquatic plants. Males guard eggs until hatching (Ref.59043).
Miller, P.J., 1986. Gobiidae. p. 1019-1085. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. Volume 3. UNESCO, Paris. (Ref. 4696)
IUCN Red List Status (Ref. 124695)
Threat to humans
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 1.0000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00708 (0.00425 - 0.01179), b=3.08 (2.94 - 3.22), in cm total length, based on LWR estimates for this species & (Sub)family-body (Ref. 93245
Trophic level (Ref. 69278
): 3.3 ±0.3 se; based on diet studies.
Resilience (Ref. 120179
): High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low to moderate vulnerability (29 of 100) .