Classification / Names
Common names | Synonyms | Catalog of Fishes(genus, species) | ITIS | CoL | WoRMS | Cloffa
Environment: milieu / climate zone / depth range / distribution range
Freshwater; benthopelagic; depth range - m (Ref. ), usually - m (Ref. ). Temperate; 0°C -
Europe: Kuban drainage from Adagum to Laba systems.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 16.0 cm SL male/unsexed; (Ref. 59043)
Morphology | Morphometrics
Diagnosed from its congeners in Black Sea basin by having the following characters: anal fin with 9½ branched rays; 36-41 + 3 scales on lateral line; interorbital distance 32-35% HL; postorbital distance 45-47% HL; eye diameter 22-26% HL, 1.3-1.5 times in snout length; body depth 21-27% SL; a row of black pigment along free margin of each flank scale, forming a regular reticulate pattern; anal and pelvic fins orange to red (Ref. 59043).
Inhabits piedmont and mountain streams with Mediterranean water regime, strong to moderate current, rock or gravel bottom. Abundant along shores and may be restricted to very small pools during summer. Feeds predominantly on terrestrial insects, benthic invertebrates and algae. Breeds in shallow riffle habitats, with strong current and stone or pebble bottom, at 10-20 cm water depth. Moves downstream in winter to deeper pools and to uppermost stretches of streams in summer (Ref. 59043).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Spawns in shallow riffles, with strong current and stone or pebble bottom, at 10-20 cm water depth. Females deposit eggs in 2-3 portions throughout the spawning period. Eggs are 1.9 in diameter, orange, adhere to pebbles and hatch in 3 days. Larvae do not possess adhesive glands, are negatively phototactic and hide under stones. They drift to shallow areas after active feeding activity commences (Ref. 59043).
Kottelat, M. and J. Freyhof, 2007. Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)
IUCN Red List Status (Ref. 124695)
Threat to humans
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00832 (0.00415 - 0.01667), b=3.13 (2.97 - 3.29), in cm total length, based on LWR estimates for this Genus-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.3 ±0.4 se; based on size and trophs of closest relatives
Resilience (Ref. 120179
): Medium, minimum population doubling time 1.4 - 4.4 years (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Moderate vulnerability (36 of 100) .