Actinopteri (ray-finned fishes) > Gobiiformes
(Gobies) > Gobiidae
(Gobies) > Gobionellinae
Etymology: Knipowitschia: Because of N.M.Knipowitsch, a zoologist and Russian ichthyologist from the Academy of Sciences. Expeditions in "Pomor" and "Pervosvanniy" ships to Azov and Black sea (1898-1908).
Environment: milieu / climate zone / depth range / distribution range
Freshwater; demersal; depth range - m (Ref. ), usually - m (Ref. ). Subtropical; 0°C -
Europe: Bosnia-Herzegovina: lower Neretva drainage. Croatia: upper Matica drainage, Neretva springs and Lake Bacina (Ref.13696, Ref.59043).
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 4.7 cm SL male/unsexed; (Ref. 59043)
Morphology | Morphometrics
This species is distinguished from its congeners by the following characters: body of breeding males with 15-18 dark irregular bars; absence of cephalic lateral line canals; body scales only under pectoral; D1 6; D2 7.5 branched rays; A 9.5 branched rays (Ref. 59043).
Occurs in clear karstic springs, lakes, slow-flowing streams and small rivers. Longevity is less than 2 years. Spawns after first winter with males defending the eggs in cavities under stones, plant material or shells. Juveniles are free swimming while adults are benthic. During summer droughts, this species may survive in underground waters (Ref.59043).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Kottelat, M., 1997. European freshwater fishes. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, 52/Suppl. 5:1-271. (Ref. 13696)
IUCN Red List Status (Ref. 124695)
Threat to humans
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.01023 (0.00444 - 0.02358), b=3.05 (2.86 - 3.24), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.4 ±0.4 se; based on size and trophs of closest relatives
Resilience (Ref. 120179
): High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low vulnerability (10 of 100) .