Actinopteri (ray-finned fishes) > Gobiiformes
(Gobies) > Gobiidae
(Gobies) > Gobionellinae
Etymology: Pomatoschistus: Greek, poma, -atos = cover, operculum + Greek, schistos = divided (Ref. 45335).
Environment: milieu / climate zone / depth range / distribution range
Marine; freshwater; brackish; demersal; amphidromous (Ref. 51243); depth range - 12 m (Ref. 35388), usually - m (Ref. ). Temperate; 8°C - 24°C (Ref. 4944); 64°N - 20°N, 19°W - 31°E
Eastern Atlantic: Norway to Morocco, including Baltic Sea (to southern Portugal) (Ref. 4696) and western Mediterranean. Also in Mauritania and the Canary Islands.
Length at first maturity / Size / Weight / Age
Maturity: Lm ?, range 3 - ? cm
Max length : 9.0 cm TL male/unsexed; (Ref. 6303); max. reported age: 2.60 years (Ref. 40230)
(total): 6 - 8;
soft rays: 7 - 10;
Vertebrae: 30 - 32. This species is distinguished from other gobies in European freshwaters by the following characters: males with conspicuous dark proximal posterior spot on first dorsal; with cephalic lateral line canals; anterior oculoscapular canal extending to snout; branched rays in second dorsal 8-10.5; anal origin below simple of first branched ray of D2; total scales in midlateral series 39-52; completely scaled postdorsal body; caudal rounded (Ref. 59043). No scales in front of the first dorsal fin (Ref. 35388); body shape is terete, slightly depressed; eyes situated dorso-laterally; reduced swim bladders (Ref. 92840).
Enters estuaries, salt marshes and pools (Ref. 4343); usually on sand or mud (Ref. 59043). Lives in sea as well as brackish waters and migrates into limnetic sections of rivers (Ref. 92840). Also rock pools where it feeds mainly on epibenthic meiofauna like harpacticoids and small amphipods (Ref. 92840); small crustaceans, worms, chironomid larvae and mites (Ref. 4696). Its burrowing habit allows it to survive low water levels and to avoid predators (Ref. 30508). Spawns after first winter, at sea, in February - September, depending on latitude; individual females spawn several times during a season. Adhesive eggs (0.9 x 0.7 mm) are deposited under or between stones, shells and aquatic plants and males guard the eggs until hatching. Juveniles enter freshwater habitats in late summer to forage (Ref. 59043). The male defends and aerates the eggs for about 9 days.
Benthic spawner (Ref. 32023).
Maugé, L.A., 1986. Gobiidae. p. 358-388. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. (Ref. 4343)
IUCN Red List Status (Ref. 124695)
Threat to humans
Fisheries: of no interest; aquarium: commercial
Estimates based on models
Preferred temperature (Ref. 115969
): 9 - 16.1, mean 10.8 °C (based on 450 cells).
Phylogenetic diversity index (Ref. 82805
= 0.5001 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00759 (0.00394 - 0.01459), b=3.08 (2.92 - 3.24), in cm total length, based on LWR estimates for this species & Genus-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.3 ±0.3 se; based on diet studies.
Generation time: 3.7 ( na - na) years. Estimated as median ln(3)/K based on 1 growth studies.
Resilience (Ref. 120179
): Medium, minimum population doubling time 1.4 - 4.4 years (K=0.29; tm=0.8; tmax=2.6; Assuming fec < 10,000).
Vulnerability (Ref. 59153
): Low to moderate vulnerability (33 of 100) .