Environment: milieu / climate zone / depth range / distribution range
Freshwater; brackish; demersal; depth range - m (Ref. ), usually - m (Ref. ). Temperate; 0°C - ; 43°N - 40°N, 22°E - 27°E
Eurasia: Northern Aegean basin from Maritza to Pinios drainages (Turkey, Greece, Bulgaria, Macedonia).
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 32.0 cm TL male/unsexed; (Ref. 1441)
Diagnosed from its congeners in Europe by the following characters: anal fin with 15-18½ (usually 16½) branched rays; back not keeled behind dorsal base; in spawning season, males with a broad black midlateral stripe, belly and check white (Ref. 59043).
Inhabits rivers and streams, usually with relatively swift current (Ref. 59043). Also occurs in lakes and lowland water courses with little current. Feeds on invertebrates and plants (Ref. 26100). Spawns among stones in large rivers (Ref. 59043). Threatened due to habitat destruction, angling and water abstraction (Ref. 26100).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Kottelat, M. and J. Freyhof, 2007. Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)
IUCN Red List Status (Ref. 124695)
Threat to humans
Fisheries: commercial; gamefish: yes
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82805
= 0.5625 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00631 (0.00391 - 0.01018), b=3.20 (3.07 - 3.33), in cm total length, based on LWR estimates for this species & (Sub)family-body (Ref. 93245
Trophic level (Ref. 69278
): 2.7 ±0.28 se; based on food items.
Resilience (Ref. 120179
): Medium, minimum population doubling time 1.4 - 4.4 years (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Moderate vulnerability (44 of 100) .