| Reference no. | Reshetnikov, Y.S., N.G. Bogutskaya, E.D. Vasil'eva, E.A. Dorofeeva, A.M. Naseka, O.A. Popova, K.A. Savvaitova, V.G. Sideleva and L.I. Sokolov, 1997 |
| Ecosystem | Ural |
| Status | native |
| Abundance | |
| Lifestage | adults |
| Regional database | |
| Remarks | Found throughout the year in lower reaches and deltas (Ref. 59043). |