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	<dc:identifier>FB-3654</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=3654</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Ovalentaria/misc</dwc:Order>
	<dwc:Family>Pomacentridae</dwc:Family>
	<dwc:Genus>Stegastes</dwc:Genus>
	<dwc:ScientificName>Stegastes variabilis (Castelnau, 1855)</dwc:ScientificName>
	<commonName xml:lang='pt'>Donzela-cacau</commonName>
	<commonName xml:lang='da'>Tofarvet jomfrufisk</commonName>
	<commonName xml:lang='en'>Cocoa damselfish</commonName>
	<commonName xml:lang='en'>Yellow demoiselle</commonName>
	<commonName xml:lang='es'>Chopa cacao</commonName>
	<commonName xml:lang='es'>Chopita cacao</commonName>
	<commonName xml:lang='es'>Jaqueta castaña</commonName>
	<commonName xml:lang='es'>Leopoldito variable</commonName>
	<commonName xml:lang=''>可可雀鯛</commonName>
	<commonName xml:lang=''>可可雀鲷</commonName>
	<commonName xml:lang=''>杂色眶锯雀鲷</commonName>
	<commonName xml:lang=''>雜色眶鋸雀鯛</commonName>
	<synonym relationship='synonym'>Eupomacentrus chrysus Bean, 1906</synonym>
	<synonym relationship='synonym'>Eupomacentrus variabilis (Castelnau, 1855)</synonym>
	<synonym relationship='misapplied name'>Pomacentrus caudalis (non Poey, 1868)</synonym>
	<synonym relationship='synonym'>Pomacentrus flaviventer Troschel, 1865</synonym>
	<synonym relationship='synonym'>Pomacentrus variabilis Castelnau, 1855</synonym>
	<dcterms:created>1991-05-01</dcterms:created>
	<dcterms:modified>2022-04-06</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33&amp;speccode=3654'>Randall, J.E.0 Food habits of reef fishes of the West Indies. Stud. Trop. Oceanogr. Miami 5:665-847. (Ref. 33)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=3654'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1673&amp;speccode=3654'>Poey, F.0 Synopsis piscium cubensium. Catalogo Razonado de los peces de la isla de Cuba. Repertor. Fisico-natural de la Isla de Cuba 2:279-484. (Ref. 1673)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1729&amp;speccode=3654'>Guitart, D.J.0 Sinopsis de los peces marinos de Cuba. Vol. III. Orden Perciformes; Suborden Percoidei. Academia de Ciencias de Cuba, Instituto de Oceanologia, La Habana. 325-608 p. (Ref. 1729)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1731&amp;speccode=3654'>Castelnau, F.L.0 Animaux nouveaux ou rares recueillis pendant l'expédition dans les parties centrales de l'Amérique de Sud, de Rio de Janeiro à Lima, et de Lima à Para; exécutée par ordre du gouvernement Français pendant les années 1843 à 1847 ... +. Exped. Amer. Sud. v. 2:i-xii + 1-112, pls. 1-50. (Ref. 1731)</reference>
	<reference isbn='0-913235-69-5 (pbk).' url='http://www.fishbase.org/references/FBRefSummary.php?id=3814&amp;speccode=3654'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 Common and scientific names of fishes from the United States and Canada. Am. Fish. Soc. Spec. Publ. (20):183 p. (Ref. 3814)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=3654'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
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	<reference isbn='3-88244-008-2.' url='http://www.fishbase.org/references/FBRefSummary.php?id=7247&amp;speccode=3654'>Allen, G.R.0 Damselfishes of the world. Mergus Publishers, Melle, Germany. 271 p. (Ref. 7247)</reference>
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	<reference isbn='0-00-219974-2 (pbk.).' url='http://www.fishbase.org/references/FBRefSummary.php?id=9710&amp;speccode=3654'>Lieske, E. and R. Myers0 Collins Pocket Guide. Coral reef fishes. Indo-Pacific &amp; Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710)</reference>
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	<reference isbn='1-56098-985-8' url='http://www.fishbase.org/references/FBRefSummary.php?id=54206&amp;speccode=3654'>Claro, R. and L.R. Parenti0 The marine ichthyofauna of Cuba. p. 21-57. In Claro, R., K.C. Lindeman and L.R. Parenti (eds) Ecology of the marine fishes of Cuba. Smithsonian Institution Press, Wahsington and London. 253p. (Ref. 54206)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78464&amp;speccode=3654'>McEachran, J.D. and J.D. Fechhelm0 Fishes of the Gulf of Mexico. Vol. 2. Austin : University of Texas Press. viii+1004p. (Ref. 78464)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=79593&amp;speccode=3654'>Albins, M.A. and M.A. Hixon0 Invasive Indo-Pacific lionfish &lt;i&gt;Pterois volitans&lt;/i&gt; reduce recruitment of Atlantic coral-reef fishes. Mar. Ecol. Prog. Ser. 367:233-238. (Ref. 79593)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=80860&amp;speccode=3654'>Menezes, N.A., P.A. Buckup, J.L. Figueiredo and R.L. Moura0 Catálogo das espécies de peixes marinhos do Brasil. São Paulo. Museu de Zoologia da Universidade de São Paulo, 160 pp. (Ref. 80860)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=82801&amp;speccode=3654'>González-Sansón, G., C. Aguilar, I. Hernández and Y. Cabrera0 Effects of depth and bottom communities on the distribution of highly territorial reef fish in the northwestern region of Cuba. J. Appl. Ichthyol. 25:652-660. (Ref. 82801)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126983&amp;speccode=3654'>IUCN0 The IUCN Red List of Threatened Species. Version 2022-2. &lt;https://www.iucnredlist.org/&gt;. Downloaded 10 Jan 2023. (Ref. 126983)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128811&amp;speccode=3654'>Harborne, A.0 Fish and coral species lists compiled by Coral Cay Conservation:  Belize 1990-1998. Coral Cay Conservation, 26p. (Ref. 128811)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=3654'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=50' logoURL='' role='photographer'>John E. Randall</agent>
	<dcterms:created>1995-01-17</dcterms:created>
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	<dc:rights>This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged</dc:rights>
	<dcterms:rightsHolder> FAO</dcterms:rightsHolder>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<dc:description>Adults inhabit inshore and offshore coral reefs.  They feed mainly on benthic algae but also on sponges, ascidians and anemones while juveniles feed on invertebrates like harpacticoid copepods and nemerteans (Ref. 9626).  Often solitary (Ref. 9626).  Juveniles are aggressively territorial (Ref. 9626).  Aggressive when breeding (Ref. 9710).  Oviparous, distinct pairing during breeding (Ref. 205).  Eggs are demersal and adhere to the substrate (Ref. 205).  Males guard and aerate the eggs (Ref. 205).  Taken incidentally in traps and small-meshed beach nets (Ref. 5217).</dc:description>
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	<reference isbn='3-88244-008-2.' url='http://www.fishbase.org/references/FBRefSummary.php?id=7247&amp;speccode=3654'>Allen, G.R. 1991 Damselfishes of the world. Mergus Publishers, Melle, Germany. 271 p. (Ref. 7247)</reference></dataObject><dataObject>
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	<dc:description>Caudal fin slightly forked, with rounded lobes.  Dark brown or blue above, shading to yellow below.  Narrow, dark, obliquely vertical lines on side.  Snout and forehead have bright blue lines.  Small dark spot in pectoral axil, another on top of caudal peduncle (Ref. 26938).</dc:description>
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	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=3654</dc:source>
	<subject></subject>
	<dc:description>Dorsal spines (total): 12; Dorsal soft rays (total): 14 - 17;  Analspines: 2; Analsoft rays: 12 - 15</dc:description>
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	<reference isbn='3-88244-008-2.' url='http://www.fishbase.org/references/FBRefSummary.php?id=7247&amp;speccode=3654'>Allen, G.R. 1991 Damselfishes of the world. Mergus Publishers, Melle, Germany. 271 p. (Ref. 7247)</reference></dataObject><dataObject>
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	<dcterms:created>1991-05-02 00:00:00</dcterms:created>
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	<dc:description>Western Atlantic:  Brazil.  Caribbean (described from Cuban specimens) species refers to Stegastes xanthurus (formerly considered as a junior synonym of S. variabilis).  Populations from Brazil and the Caribbean are genetically very distinct (Roberstson &amp; van Tassell, 2012; accessed 08 August 2016; http://biogeodb.stri.si.edu/caribbean/en/thefishes/species/3884).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95977&amp;speccode=3654'>Smith-Vaniz, W.F. and H.L. Jelks 2014 Marine and inland fishes of St. Croix, U.S. Virgin Islands: an annotated checklist. Zootaxa 3803(1):1-120. (Ref. 95977)</reference></dataObject><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2022-04-06 00:00:00</dcterms:modified>
	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>fisheries: commercial; aquarium: commercial</dc:description>
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	<reference isbn='3-88244-008-2.' url='http://www.fishbase.org/references/FBRefSummary.php?id=7247&amp;speccode=3654'>Allen, G.R. 1991 Damselfishes of the world. Mergus Publishers, Melle, Germany. 271 p. (Ref. 7247)</reference>
	<reference isbn='3-88244-008-2.' url='http://www.fishbase.org/references/FBRefSummary.php?id=7247&amp;speccode=3654'>Allen, G.R. 1991 Damselfishes of the world. Mergus Publishers, Melle, Germany. 271 p. (Ref. 7247)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=3654&amp;StockCode=3850</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Oviparous, distinct pairing during breeding (Ref. 205).  Eggs are demersal and adhere to the substrate (Ref. 205).  Males guard and aerate the eggs (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=3654'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-3654</dc:identifier>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2022-04-06 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses</subject>
	<dc:description>12.5 cm TL (male/unsexed; (Ref. 9626))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9626'>Cervigón, F. 1993 Los peces marinos de Venezuela. Volume 2. Fundación Científica Los Roques, Caracas,Venezuela. 497 p.</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2022-04-06 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=3654</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>reef-associated; non-migratory; marine; depth range 0 - 30 m (Ref. 7247)</dc:description>
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	<reference isbn='3-88244-008-2.' url='http://www.fishbase.org/references/FBRefSummary.php?id=7247&amp;speccode=3654'>Allen, G.R. 1991 Damselfishes of the world. Mergus Publishers, Melle, Germany. 271 p. (Ref. 7247)</reference></dataObject>
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