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	<dc:identifier>FB-4062</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4062</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Cottoidei</dwc:Order>
	<dwc:Family>Cottidae</dwc:Family>
	<dwc:Genus>Cottus</dwc:Genus>
	<dwc:ScientificName>Cottus asper Richardson, 1836</dwc:ScientificName>
	<commonName xml:lang='da'>Pigget ulk</commonName>
	<commonName xml:lang='en'>Bullhead</commonName>
	<commonName xml:lang='en'>Prickly bullhead</commonName>
	<commonName xml:lang='en'>Prickly sculpin</commonName>
	<commonName xml:lang='fr'>Chabot piquant</commonName>
	<commonName xml:lang=''>Hl'aama</commonName>
	<commonName xml:lang=''>多棘杜父魚</commonName>
	<commonName xml:lang=''>多棘杜父鱼</commonName>
	<synonym relationship='synonym'>Centridermichthys asper (Richardson, 1836)</synonym>
	<synonym relationship='synonym'>Centridermichthys parvus (Girard, 1854)</synonym>
	<synonym relationship='synonym'>Cottopsis asper (Richardson, 1836)</synonym>
	<synonym relationship='synonym'>Cottopsis parvus Girard, 1854</synonym>
	<synonym relationship='synonym'>Trachidermus richardsonii Heckel, 1837</synonym>
	<dcterms:created>1991-05-01</dcterms:created>
	<dcterms:modified>1999-11-19</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=265&amp;speccode=4062'>Matarese, A.C., A.W. Kendall, D.M. Blood and M.V. Vinter0 Laboratory guide to early life history stages of Northeast Pacific fishes. NOAA Tech. Rep. NMFS 80:1-652. (Ref. 265)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=276&amp;speccode=4062'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 A list of common and scientific names of fishes from the United States and Canada. Am. Fish. Soc. Spec. Publ. (12)1-174. (Ref. 276)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1998&amp;speccode=4062'>Scott, W.B. and E.J. Crossman0 Freshwater fishes of Canada. Bull. Fish. Res. Board Can. 184: xi+1-966. (Ref. 1998)</reference>
	<reference isbn='0-395-26873-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=2850&amp;speccode=4062'>Eschmeyer, W.N., E.S. Herald and H. Hammann0 A field guide to Pacific coast fishes of North America. Boston (MA, USA): Houghton Mifflin Company. xii+336 p. (Ref. 2850)</reference>
	<reference isbn='0-913235-69-5 (pbk).' url='http://www.fishbase.org/references/FBRefSummary.php?id=3814&amp;speccode=4062'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 Common and scientific names of fishes from the United States and Canada. Am. Fish. Soc. Spec. Publ. (20):183 p. (Ref. 3814)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=4062'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=4062'>Page, L.M. and B.M. Burr0 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6876&amp;speccode=4062'>Quast, J.C. and E.L. Hall0 List of fishes of Alaska and adjacent waters with a guide to some of their literature. U.S. Dep. Commer., NOAA Tech. Rep. NMFS SSRF-658, 47 p. (Ref. 6876)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10023&amp;speccode=4062'>Richardson, J.0 The fish. Fauna Boreali-Americana; or the zoology of of the northern parts of British America. (3):327 p. (Ref. 10023)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10280&amp;speccode=4062'>McPhail, J.D. and C.C. Lindsey0 Freshwater fishes of northwestern Canada and Alaska. Fish. Res. Board Can. Bull. 173:381 p. (Ref. 10280)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10354&amp;speccode=4062'>Heckel, J.J.0 Ichthyologische Beitrage zu den Familien der Cottoiden, Scorpaenoiden, gobioiden, und Cyprinoiden. Ann. Wein. Mus. Natur. 2:143-164. (Ref. 10354)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10355&amp;speccode=4062'>Girard, C.F.0 Contributions to the natural history of the freshwater fishes of North America. I. A monograph of the cottoids. Smithsonian Contrib. Knowl. 3(art. 3):80 p. (Ref. 10355)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10356&amp;speccode=4062'>Lord, J.K.0 The naturalist in Vancouver Island and British Columbia. Vol. 2. Richard Bentley, London. 375 p. (Ref. 10356)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10357&amp;speccode=4062'>Munro, J.A. and W.A. Clemens0 Food of the American merganser &lt;i&gt;Mergus merganser americanus&lt;/i&gt; in British Columbia. Can. Field-Natur. 50(3):34-36. (Ref. 10357)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10882&amp;speccode=4062'>McPhail, J.D. and R. Carveth0 Field key to the freshwater fishes of British Columbia. Fish Museum, Department of Zoology, U.B.C., Canada, 239 p. (Ref. 10882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10937&amp;speccode=4062'>Anonymous0 Fish collection database of the University of British Columbia Fish Museum Fish Museum. University of British Columbia, Vancouver, Canada. (Ref. 10937)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11366&amp;speccode=4062'>Evermann, B.W. and E.L. Goldsborough0 The fishes of Alaska. Bull. U.S. Bur. Fish. 26: 219-360. (Ref. 11366)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12204&amp;speccode=4062'>Coad, B.W.0 Encyclopedia of Canadian fishes. Canadian Museum of Nature and Canadian Sportfishing Productions Inc. Singapore. (Ref. 12204)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13348&amp;speccode=4062'>Brown, L.R., S.A. Matern and P.B. Moyle0 Comparative ecology of prickly sculpin, &lt;i&gt;Cottus asper&lt;/i&gt;, and coastrange sculpin, &lt;i&gt;C. aleuticus&lt;/i&gt;, in the Eel River, California. Environ. Biol. Fishes 42:329-343. (Ref. 13348)</reference>
	<reference isbn='0-7099-5503-0' url='http://www.fishbase.org/references/FBRefSummary.php?id=26213&amp;speccode=4062'>McDowall, R.M.0 Diadromy in fishes: migrations between freshwater and marine environments. Croom Helm, London. (Ref. 26213)</reference>
	<reference isbn='0-920080-75-8.' url='http://www.fishbase.org/references/FBRefSummary.php?id=27436&amp;speccode=4062'>Lamb, A. and P. Edgell0 Coastal fishes of the Pacific northwest. Madeira Park, (BC, Canada): Harbour Publishing Co. Ltd., 224 p. (Ref. 27436)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27547&amp;speccode=4062'>Morrow, J.E.0 The freshwater fishes of Alaska. University of. B.C. Animal Resources Ecology Library. 248p. (Ref. 27547)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28210&amp;speccode=4062'>Bond, C.E.0 Distribution and ecology of freshwater sculpins, genus &lt;i&gt;Cottus&lt;/i&gt;, in Oregon. Univ. Michigan, Ann Arbor. 186 pp. PhD thesis. (Ref. 28210)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28919&amp;speccode=4062'>Krejsa, R.J.0 The systematics of the prickly sculpin, &lt;i&gt;Cottus asper&lt;/i&gt; Richardson, a polytypic species. Part I. Synonymy, nomenclatural history, and distribution. Pac. Sci. 21(2):241-251. (Ref. 28919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28920&amp;speccode=4062'>Sinclair, D.C.0 Diel limnetic occurrence of young &lt;i&gt;Cottus asper&lt;/i&gt; in two British Columbia lakes. J. Fish. Res. Board Can. 25(9):1997-2000. (Ref. 28920)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=4062'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58470&amp;speccode=4062'>Wydoski, R.S. and R.R. Whitney0 Inland fishes of Washington. University Washington Press. 220p. (Ref. 58470)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=4062'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=4062'>Corush, J.B.0 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-18 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>There appears to be two genetically distinct forms (Ref. 27547), an inland form found in sandy and rocky shores of lakes, and a coastal form usually found over sand in quiet runs of small to medium rivers; sometimes in salt water near river mouths (Ref. 2850).  The coastal form moves into brackish estuaries to spawn (Ref. 27547).  Oviparous with demersal, adhesive eggs and pelagic larvae (Ref. 265).  Adults feed mainly on aquatic insect larvae and bottom invertebrates (Ref. 1998).  Too small to be used as food and too difficult to capture in large numbers to be used for anything else (Ref. 27547) but large individuals are reported to be excellent eating as well as good bait fishes (Ref. 2850).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=4062'>Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
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	<dc:description>Distinguished by a complete lateral line, a single pore at the tip of the chin, the presence of 15 to 19 anal rays, and well developed palatine teeth (Ref. 27547).  Pectorals large and fan-shaped; caudal truncate or slightly rounded (Ref. 27547).  Dark brown, olive or gray above and on sides, whitish yellow to white below; usually three dark irregular blotches or bars below soft dorsal; vague irregular dark mark on sides; fins (except anal) have dark bars, the first dorsal with a dark spot towards the rear (Ref. 27547).  Both sexes show an orange band on the edge of the first dorsal fin at spawning time (Ref. 27547).  The inland form is generally more densely prickled over a larger portion of the body while the coastal form shows a reduced number of prickles (Ref. 28211).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27547&amp;speccode=4062'>Morrow, J.E. 1980 The freshwater fishes of Alaska. University of. B.C. Animal Resources Ecology Library. 248p. (Ref. 27547)</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 7 - 10; Dorsal soft rays (total): 18 - 23;  Analspines: 0; Analsoft rays: 15 - 19; Vertebrae: 34 - 39</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27547&amp;speccode=4062'>Morrow, J.E. 1980 The freshwater fishes of Alaska. University of. B.C. Animal Resources Ecology Library. 248p. (Ref. 27547)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-4062</dc:identifier>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-02 00:00:00</dcterms:created>
	<dcterms:modified>2024-12-05 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4062</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>North America:  Pacific slope drainages from Seward, Alaska to Ventura River, California, USA; also east of Continental Divide in upper Peace River in British Columbia, Canada.  Exhibits coastal and inland forms that are genetically distinct (Ref. 27547).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=4062'>Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-4062</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-18 00:00:00</dcterms:modified>
	<dc:title>Importance</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4062</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>fisheries: commercial; aquarium: public aquariums; bait: occasionally</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9183'>Newman, L. 1995 Census of fish at the Vancouver aquarium, 1994. Unpublished manuscript.</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9183'>Newman, L. 1995 Census of fish at the Vancouver aquarium, 1994. Unpublished manuscript.</reference>
	<reference isbn='0-395-26873-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=2850&amp;speccode=4062'>Eschmeyer, W.N., E.S. Herald and H. Hammann 1983 A field guide to Pacific coast fishes of North America. Boston (MA, USA): Houghton Mifflin Company. xii+336 p. (Ref. 2850)</reference></dataObject><dataObject>
	<dc:identifier>FB-Migration-4062</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-18 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4062</dc:source>
	<dc:description>Catadromous. Migrating from freshwater to the sea to spawn, e.g., European eels.  Subdivision of diadromous.  Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=4062'>Corush, J.B. 2019 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference></dataObject><dataObject>
	<dc:identifier>FB-LifeCycle-4062</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-18 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=4062&amp;StockCode=4258</dc:source>
	<dc:description>Prior to breeding, males move downstream and select  a nesting site under boulders or flat rocks. Females aggregate some distance upstream and move down singly to the spawning area. Courtship behavior occurs outside the nest until a  female is selected. The pair enter the nesting site, courting continues until eggs are deposited and fertilized. The female then leaves the nest and goes back upstream to feed, while the male spawns with other females or fans and guards the eggs. The male does not feed until the eggs have hatched, moving upstream only in the late summer (Ref. 1998, 27547). During the planktonic stage, larvae of the freshwater nonanadromous form in lakes show distinct diurnal vertical migrations, being most abundant at the surface during the darkest hours of the night. They apparently stay deep in the water during the day and on bright moonlight nights (Ref. 28920). Metamorphosis is complete by the end of the planktonic period and the young take up a demersal mode of life. The young coastal form may move upstream during the fall, although the young may remain in the estuary for a full year (Ref. 27547).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1998&amp;speccode=4062'>Scott, W.B. and E.J. Crossman 1973 Freshwater fishes of Canada. Bull. Fish. Res. Board Can. 184: xi+1-966. (Ref. 1998)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-4062</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1997-09-29 00:00:00</dcterms:created>
	<dcterms:modified>2008-04-03 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>asper</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>There appears to be two genetically distinct forms (Ref. 27547), an inland form found in sandy and rocky shores of lakes, and a coastal form usually found over sand in quiet runs of small to medium rivers; sometimes in salt water near river mouths (Ref. 2850).  The coastal form moves into brackish estuaries to spawn (Ref. 27547).   Feed mainly on aquatic insect larvae and bottom invertebrates (Ref. 1998). Juvenile and adults feed mainly on insects (Ref. 13348).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13348&amp;speccode=4062'>Brown, L.R., S.A. Matern and P.B. Moyle 1995 Comparative ecology of prickly sculpin, &lt;i&gt;Cottus asper&lt;/i&gt;, and coastrange sculpin, &lt;i&gt;C. aleuticus&lt;/i&gt;, in the Eel River, California. Environ. Biol. Fishes 42:329-343. (Ref. 13348)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-4062</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-18 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4062</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>30.0 cm TL (male/unsexed; (Ref. 5723)); max. reported age: 7 years (Ref. 28210)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=4062'>Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28210&amp;speccode=4062'>Bond, C.E. 1963 Distribution and ecology of freshwater sculpins, genus &lt;i&gt;Cottus&lt;/i&gt;, in Oregon. Univ. Michigan, Ann Arbor. 186 pp. PhD thesis. (Ref. 28210)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder> FAO</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject></subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-4062</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-18 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4062</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>demersal; catadromous (Ref. 138280); freshwater; brackish; marine</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=4062'>Corush, J.B. 2019 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference></dataObject>
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