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	<dc:identifier>FB-52470</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=52470</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Blenniiformes</dwc:Order>
	<dwc:Family>Clinidae</dwc:Family>
	<dwc:Genus>Heteroclinus</dwc:Genus>
	<dwc:ScientificName>Heteroclinus johnstoni (Saville-Kent, 1886)</dwc:ScientificName>
	<commonName xml:lang='en'>Witch guitarfish</commonName>
	<commonName xml:lang='et'>Kare kitarrrai</commonName>
	<commonName xml:lang='es'>Guitarra bruja</commonName>
	<commonName xml:lang=''>光滑強鰭鰩</commonName>
	<commonName xml:lang=''>光滑强鳍鳐</commonName>
	<dcterms:created>1999-05-05</dcterms:created>
	<dcterms:modified>2014-01-10</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=52476'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10937&amp;speccode=52476'>Anonymous0 Fish collection database of the University of British Columbia Fish Museum Fish Museum. University of British Columbia, Vancouver, Canada. (Ref. 10937)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13259&amp;speccode=52476'>Acero, A.P. and R. Franke0 Nuevos registros de peces cartilaginosos del Parque Nacional Natural Gorgona (Pacífico Colombiano), II. Rayas y descripción de una nueva especie. Biblioteca Jose Jeronimo Triana (11):9-21. (Ref. 13259)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26282&amp;speccode=52476'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p. (Ref. 26282)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=52476'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=52476'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=52476'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=52476'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=96293&amp;speccode=52476'>Page, L.M., H. Espinosa-Pérez, L.T. Findley, C.R. Gilbert, R.N. Lea, N.E. Mandrak, R.L. Mayden and J.S. Nelson0 Common and scientific names of fishes from the United States, Canada and Mexico, 7th edition. Bethesda, Maryland: American Fisheries Society, Special Publication 34, 384 p. (Ref. 96293)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=97056&amp;speccode=52476'>Robertson, D. R. and G.R. Allen0 Shorefishes of the tropical eastern Pacific online information system. Version 1.0. Smithsonian Tropical Research Institute, Balboa, Panama. Available: www.stri.org/sftep. (Ref. 97056)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105120&amp;speccode=52476'>Espinoza, M., T.M. Clarke, F. Villalobos-Rojas and I.S. Wehrtmann0 Diet composition and diel feeding behaviour of the banded guitarfish &lt;i&gt;Zapteryx xyster&lt;/i&gt; along the Pacific coast of Costa Rica, Central America. J. Fish Biol. 82(1):286-305. (Ref. 105120)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=52476'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=52476'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor0 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123526&amp;speccode=52476'>Garcés, H.0 Lista sistemática preliminar de los peces marinos comerciales del Pacifico de Panama. Tecnociencia, 23(1):198-237. (Ref. 123526)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=127662&amp;speccode=52476'>Kyne, P.M., A.J. Courtney, I.P. Jacobsen and M.B. Bennett0 Reproductive parameters of rhinobatid and urolophid batoids taken as by‐catch in the Queensland (Australia) east coast otter‐trawl fishery. J. Fish Biol. 89(2):1208-1226. (Ref. 127662)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=127741&amp;speccode=52476'>Clarke, T.M., M. Espinoza and I.S. Wehrtmann0 Reproductive ecology of demersal elasmobranchs from a data-deficient fishery, Pacific of Costa Rica, Central America. Fisheries Research 157:96-105. (Ref. 127741)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130539&amp;speccode=52476'>Zavalaga Talledo, F., S.V. Campos León and K. Kanagusuku Gondo0 Guía para la determinación de las principales rayas del Perú. Lima (Peru): Instituto del Mar del Perú (IMARPE), 42 p. (Ref. 130539)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-52476</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-05-05 00:00:00</dcterms:created>
	<dcterms:modified>2024-08-16 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<audience>General public</audience>
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	<dc:description>Benthic species of continental shelf on rocky and sandy bottoms to depths of 150 m.  It is a night feeder, mainly on small fishes and prawns (Ref. 114953).  Ovoviviparous (Ref. 50449).  Maturity size of males at ca. 47 cm TL, females at ca. 45 cm TL. Species born at ca. 18 cm TL.  It is a likely bycatch of gillnet and prawn trawl fisheries (Ref. 114953).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13259&amp;speccode=52476'>Acero, A.P. and R. Franke 1995 Nuevos registros de peces cartilaginosos del Parque Nacional Natural Gorgona (Pacífico Colombiano), II. Rayas y descripción de una nueva especie. Biblioteca Jose Jeronimo Triana (11):9-21. (Ref. 13259)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-52476</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-05-05 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=52476</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Eastern Pacific: southeastern Gulf of California, Mexico to Peru (Ref. 97056). Not recorded in Compagno's 1999 checklist (Ref. 35766).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13259&amp;speccode=52476'>Acero, A.P. and R. Franke 1995 Nuevos registros de peces cartilaginosos del Parque Nacional Natural Gorgona (Pacífico Colombiano), II. Rayas y descripción de una nueva especie. Biblioteca Jose Jeronimo Triana (11):9-21. (Ref. 13259)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-52476</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-05-05 00:00:00</dcterms:created>
	<dcterms:modified>2024-08-16 00:00:00</dcterms:modified>
	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>fisheries: bycatch</dc:description>
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	<dc:identifier>FB-LifeCycle-52476</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1999-05-05 00:00:00</dcterms:created>
	<dcterms:modified>2024-08-16 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=52476&amp;StockCode=43652</dc:source>
	<dc:description>Exhibit ovoviparity (aplacental viviparity), with embryos feeding initially on yolk, then receiving additional nourishment from the mother by indirect absorption of uterine fluid enriched with mucus, fat or protein through specialised structures (Ref. 50449).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=52476'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-52476</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
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	<dcterms:modified>2024-08-16 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=52476</dc:source>
	<dc:description>78.0 cm TL (male/unsexed; (Ref. 114953))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=52476'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor 2016 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-52476</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-05-05 00:00:00</dcterms:created>
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	<dc:title>Environment</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>reef-associated; marine; depth range ? - 150 m (Ref. 114953)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=52476'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor 2016 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference></dataObject>
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