FB-5525 http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 Animalia Chordata Teleostei Perciformes/Serranoidei Epinephelidae Epinephelus Epinephelus tukula Morgans, 1959 Garoupa batata Aartappel-baars Kartoffelbars Black-spotted rock-cod Grouper Potato bass Potato cod Potato grouper Potato rockcod Mérou patate Lokos Inid orig Kasurihata Sumeyn Mero patata Cabot malgache Garea Hassine batata Kiapu buntar tikolo Lilla Maleog Okke Vualia 石斑 蓝身大斑石斑鱼 藍身大斑石斑魚 藍身大石斑魚 过鱼 金錢躉 鲙 Epinephelus fuscoguttatus (non Forsskål, 1775) Serranus dispar var. a Playfair, 1867 Serranus fuscoguttatus (non Forsskål, 1775) 1992-06-02 2013-10-22 Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171) Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino0 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559) Smith, J.L.B. and M.M. Smith0 The fishes of Seychelles. Rhodes University, Grahamstown. 215 p. (Ref. 1623) Eschmeyer, W.N.0 Catalog of the genera of recent fishes. California Academy of Sciences, San Francisco, USA. 697 p. (Ref. 1830) Randall, J.E., G.R. Allen and R.C. Steene0 Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, Hawaii. 506 p. (Ref. 2334) Bianchi, G.0 FAO species identification sheets for fishery purposes. Field guide to the commercial marine and brackish-water species of Pakistan. Prepared with the support of PAK/77/033 and FAO (FIRM) Regular Programme. Rome: FAO. 200 p. (Ref. 2872) Heemstra, P.C. and J.E. Randall0 Serranidae. In W. Fischer and G. Bianchi (eds.). FAO species identification sheets for fishery purposes. Western Indian Ocean (fishing area 51). Rome, FAO. Vol. IV. 241 p. (Ref. 3153) van der Elst, R.0 A guide to the common sea fishes of southern Africa. C. Struik, Cape Town. 367 p. (Ref. 3670) Torres, F.S.B. Jr.0 Tabular data on marine fishes from Southern Africa, Part I. Length-weight relationships. Fishbyte 9(1):50-53. (Ref. 3678) Smith, A. and P. Dalzell0 Fisheries resources and management investigations in Woleai Atoll, Yap State, Federated States of Micronesia. Inshore Fish. Res. Proj., Tech. Doc., South Pacific Commission. Noumea, New Caledonia. 64 p. (Ref. 3807) Heemstra, P.C. and J.E. Randall0 Serranidae. p. 509-537. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 4319) Randall, J.E. and P.C. Heemstra0 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p. (Ref. 4787) Shao, K.-T., S.-C. Shen, T.-S. Chiu and C.-S. Tzeng0 Distribution and database of fishes in Taiwan. p. 173-206. In C.-Y. Peng (ed.) Collections of research studies on 'Survey of Taiwan biological resources and information management'. 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Öhman0 Coral and fish distribution patterns in Mafia Island Marine Park, Tanzania: fish-habitat interactions. Hydrobiologia 498:191-211. (Ref. 51015) Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471) Letourneur, Y., P. Chabanet, P. Durville, M. Taquet, E. Teissier, M. Parmentier, J.-C. Quéro and K. Pothin0 An updated checklist of the marine fish fauna of Reunion Island, south-western Indian Ocean. Cybium 28(3):199-216. (Ref. 53568) Chen, C.-H.0 Checklist of the fishes of Penghu. FRI Special Publication No. 4. 175 p. (Ref. 55073) Pereira, M.A.M.0 Preliminary checklist of reef-associated fishes of Mozambique. Maputo, Ministtry for the Coordination of Environmental Affairs (MICOA). 21 pp. (Ref. 57683) Baissac, J. de B.0 SWIOP/WP/54 - Checklist of the marine fishes of Mauritius. RAF/87/008/WP/54/90 Regional Project for the Development & Management of Fisheries in the Southwest Indian Ocean. 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(Ref. 83882) Golani, D. and S. V. Bogorodsky0 The fishes of the Red Sea - reappraisal and updated checklist. Zootaxa 2463:1-135. (Ref. 84159) May, D.0 Folk taxonomy of reef fish and the value of participatory monitoring in Wakatobi National Park, southeast Sulawesi, Indonesia. SPC Traditional Marine Resource Management and Knowledge Information Bulletin 18: 18-35. (Ref. 87848) Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra0 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group, xix, 356 p., A47 pages appendix. (Ref. 89707) Taquet, M. and A. Diringer0 Mérous de l'Océan Indien. La Réunion: Azalées Editions. 103 p. (Ref. 89967) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062) Allen, G.R. and M.V. Erdmann0 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. 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(Ref. 101110) Koror Republic of Palau.0 The Rock Islands Southern Lagoon. http://www.google.com/url?sa=t&rct=j&q=&esrc=s&source=web&cd=3&cad=rja&uact=8&ved=0CCkQFjAC&url=http%3A%2F%2Fwww.palaupanfund.org%2Fpdf%2Fmanagementplan%2Fkoror%2FRock%2520Islands%2520Southern%2520Lagoon%2520Area%2520Management%2520Plan%25202012-2016%2520latest%2520July%252018.pdf&ei=HKOcVZ6LMNjr8AXEuKzwCA&usg=AFQjCNFN07tAwxC1s67J47zoXSQImcfNmg&sig2=z7WLKekW7DHqs_11pn39ZA&bvm=bv.96952980,d.dGc [Accessed 08/07/2015]. (Ref. 103126) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272) SCRFA, Science and Conservation of Fish Aggregations0 SCRFA database. Web online database. https://www.scrfa.org/database/: SCRFA. [2013-onwards]. (Ref. 118580) Allen, G.R. and M.V. Erdmann0 Coral reef fishes of Timor-Leste. p. 33-82. In G.R. Allen and M.V. Erdmann (eds). A rapid biological assessment of Timor-Leste. Conservation International. (Ref. 118793) University of Western Australia0 Benthic BRUVS data 2019. Video BRUVS from Jessica Meeuwig. (Ref. 122752) van der Elst, Rudy, Simon Chater and Dennis King0 Fishes from Bassas Da India Atoll in the Mozambique Channel. Atoll Research Bulletin No. 629. (Ref. 124899) Lai, N.-W.0 Hong Kong market marine fish. Original title: 香港街市海魚圖鑑. Hua Qianshu Publishing Co., Ltd., p. 440. (Ref. 129440) Béarez, P. and L. Bouffandeau0 Fishing for survival: The forgotten slaves of Tromelin Island (Indian Ocean). Int J Osteoarchaeol. 29:487-498. (Ref. 137464) Petit-Marty, N., M. Vázquez-Luis and I.E. Hendriks0 Use of the nucleotide diversity in COI mitochondrial gene as an early diagnostic of conservation status of animal species. Conserv. Lett. 14(1):1-7. (Ref. 138103) FB-pic-5525-7345 http://purl.org/dc/dcmitype/StillImage image/gif FAO 1996-01-15 1996-04-19 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution http://www.fishbase.us/images/species/Eptuk_u0.gif http://www.fishbase.us/images/thumbnails/gif/tn_Eptuk_u0.gif FB-pic-5525-9646 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-03-04 1999-07-07 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://www.fishbase.us/images/species/Eptuk_u1.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Eptuk_u1.jpg FB-pic-5525-13431 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1997-02-19 2011-05-17 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size http://www.fishbase.us/images/species/Eptuk_u0.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Eptuk_u0.jpg FB-GeneralDescr-5525 http://purl.org/dc/dcmitype/Text text/html Crispina B. Binohlan 1992-06-02 00:00:00 2013-10-22 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Mainly found in deep reef channels and seamounts, in current prone areas (Ref. 48635). Juveniles may be found in tide pools (Ref. 5222). Feeds on reef fishes, skates, crabs, and spiny lobsters (Ref. 5222). Considered to be exceedingly territorial and very aggressive towards intruders. Vulnerable to spear fishers (Ref. 5222). Hand fed by divers in certain areas, but potentially dangerous to the inexperienced. In the Hong Kong live fish markets (Ref. 27253). Solitary (Ref 90102). Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-DiagnosticDescr-5525 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-06-15 00:00:00 2013-10-22 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Distinguished by the following characteristics: body depth contained 2.9-3.5 times in SL; head length 2.3-2.6 times in SL; slightly convex interorbital area; straight dorsal head profile; rounded or subangular preopercle, slightly enlarged serrae at angle; almost straight upper edge of operculum; posterior and anterior nostrils subequal in size; maxilla reaches to or beyond vertical at rear edge of eye; 2-6 rows of teeth on midlateral part of lower jaw (Ref. 89707). Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Morphology-5525 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-06-15 00:00:00 2013-10-22 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription Dorsal spines (total): 11; Dorsal soft rays (total): 14 - 15; Analspines: 3; Analsoft rays: 8 Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Distribution-5525 http://purl.org/dc/dcmitype/Text text/html Crispina B. Binohlan 1992-06-02 00:00:00 2005-10-26 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology Indo-West Pacific: Red Sea and East Africa to southern Japan and Queensland, Australia. Also from the Paracel Islands in the South China Sea. There are no records from Madagascar, Mauritius, Maldives, Laccadives, and Sri Lanka, Indonesia, and the Philippines. Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Uses-5525 http://purl.org/dc/dcmitype/Text text/html Crispina B. Binohlan 1992-06-02 00:00:00 2013-10-22 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution fisheries: subsistence fisheries; gamefish: yes Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter 1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171) Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-TrophicStrategy-5525 http://purl.org/dc/dcmitype/Text text/html Drina Sta. Iglesia 2000-11-13 00:00:00 2024-02-06 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ tukula Dwells in coral and rocky reefs. Solitary, but forms pairs when spawning (Ref. 9773). Mainly found in deep reef channels and seamounts, in current prone areas (Ref. 48635). Juveniles may be found in tide pools (Ref. 5222). Also in the external areas around coral reefs (Ref. 9137). Feed on reef fishes, skates, crabs, and spiny lobsters (Refs. 5222, 9773). Considered to be exceedingly territorial and very aggressive towards intruders (Also Ref. 9773). Vulnerable to spear fishers (Ref. 5222). Large adults would be potential predators of small Latimeria (Ref. 58472). Hand fed by divers in certain areas, but potentially dangerous to the inexperienced. Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Size-5525 http://purl.org/dc/dcmitype/Text text/html Crispina B. Binohlan 1992-06-02 00:00:00 2013-10-22 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy 200 cm TL (male/unsexed; (Ref. 2872)); max. published weight: 110.0 kg (Ref. 9710) Bianchi, G. 1985 FAO species identification sheets for fishery purposes. Field guide to the commercial marine and brackish-water species of Pakistan. Prepared with the support of PAK/77/033 and FAO (FIRM) Regular Programme. Rome: FAO. 200 p. (Ref. 2872) Lieske, E. and R. Myers 1994 Collins Pocket Guide. Coral reef fishes. Indo-Pacific & Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=2 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-5525 http://purl.org/dc/dcmitype/Text text/html Crispina B. Binohlan 1992-06-02 00:00:00 2013-10-22 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5525 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats reef-associated; marine; depth range 10 - 400 m (Ref. 89707) Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra 2011 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group, xix, 356 p., A47 pages appendix. (Ref. 89707)