FB-69269 http://www.fishbase.org/summary/SpeciesSummary.php?id=69269 Animalia Chordata Teleostei Anguilliformes Ophichthidae Neenchelys Neenchelys nudiceps Tashiro, Hibino & Imamura, 2015 Variedspine goatfish Varied-spine goatfish Upeneus australiae (non e Kim & Nakaya, 2002) Upeneus bensasi (non Temminck & Schlegel, 1843) Upeneus guttatus (non Day, 1868) Upeneus margarethae (non Uiblein & Heemstra, 2010) Upeneus sp. 1 Not applicable Upeneus spottocaudalis (non Uiblein & Gledhill, 2017) 2020-11-06 2020-11-11 Uiblein, F. and P.C. Heemstra0 A taxonomic review of the Western Indian Ocean goatfishes of the genus <i>Upeneus</i> (Family Mullidae), with descriptions of four new species. Smithiana 11:35-71. (Ref. 83903) Uiblein, F. and M. Lisher0 A new goatfish of the genus <i>Upeneus</i> (Mullidae) from Angoche, northern Mozambique. Zootaxa 3717(1):85-95. (Ref. 94148) Patzner, R.A.0 Reproductive strategies of fish. pp. 311-350. In Rocha, M.J., A. Arukwe and B.G. Kapoor (eds). Fish reproduction: cytology, biology and ecology. Science Publisher, Inc. Oxford. 631 p. (Ref. 101194) White, W.T., P.R. Last, Dharmadi, R. Faizah, U. Chodrijah, B.I. Prisantoso, J.J. Pogonoski, M. Puckridge and S.J.M. Blaber0 Market fishes of Indonesia (= Jenis-jenis ikan di Indonesia). ACIAR Monograph No. 155. Australian Centre for International Agricultural Research: Canberra, 438 p. (Ref. 117228) Uiblein, F., D.C. Gledhill, D.A. Pavlov, T.A. Hoang and S. Shaheen0 Three new goatfishes of the genus <i>Upeneus</i> (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in <i>U. margarethae</i>. Zootaxa 4683(2):151-196. (Ref. 121834) Motomura, H., U.B. Alama, N. Muto, P. Babaran and S. Ishikawa (eds.)0 Commercial and bycatch market fishes of Panay Island, Republic of the Philippines. The Kagoshima University Museum, Kagoshima, University of the Philippines Visayas, Iloilo, and Research Institute for Humanity and Nature, Kyoto. 246 pp, 911 figs. (Ref. 123102) FB-GeneralDescr-69278 http://purl.org/dc/dcmitype/Text text/html Nicolas Bailly 2020-11-09 00:00:00 2021-01-22 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=69278 Occurs on sandy or muddy bottoms (Ref. 121834). Uiblein, F., D.C. Gledhill, D.A. Pavlov, T.A. Hoang and S. Shaheen 2019 Three new goatfishes of the genus <i>Upeneus</i> (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in <i>U. margarethae</i>. Zootaxa 4683(2):151-196. (Ref. 121834) FB-DiagnosticDescr-69278 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 2021-01-11 00:00:00 2021-01-11 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=69278 This species is distinguished by the following characters: D VII or VIII + 9, with the first spine minute in 8-spined fish; pectoral fins 13-15; gill rakers 4-6 + 15-18 = 21-24; lateral-line scales 28-30. All measurements in % SL, adults: body depth at first dorsal-fin origin 22-26; body depth at anus 18-22; caudal-peduncle depth 9.2-11; caudal-peduncle width 3.5-5.0; maximum head depth 19-23; head depth through eye 15-18; suborbital depth 8.3-11; interorbital length 7.2-9.2; head length 27-31; snout length 9.7-13; postorbital length 10-13; orbit length 5.9-8.3; upper-jaw length 9.4-13; barbel length 16-20; interdorsal distance 13-17; caudal-peduncle length 22-26; caudal-fin length 27-32; anal-fin height 15-19; pelvic-fin length 19-23; pectoral-fin length 19-22; pectoral-fin width 3.4-4.7; first dorsal-fin height 18­­-23; second dorsal-fin height 16-20. All measurements in % SL, subadults: body depth at first dorsal-fin origin 22-24; body depthat anus 18-21; caudal-peduncle depth 9.1-10; caudal-peduncle width 3.6-3.8; maximum head depth 19-21; head depth through eye 15-18; suborbital depth 9.0-10; interorbital length 7.6-8.1; head length 29-30; snout length 11-12; postorbital length 11-13; orbit length 7.8-7.9; upper-jaw length 9.9-12; barbel length 18-19; interdorsal distance 12-14; caudal-peduncle length 22-25; caudal-fin length 29-31; anal-fin height 16-19; pelvic-fin length 21-22; pectoral-fin length 21; pectoral-fin width 3.8-4.2; first dorsal-fin height 21­-22; second dorsal-fin height 17-20. Colouration: head and body are red dorsally, brown or dark-grey mottled, ventrally white, often with pale beige or grey dots or red blotches (inactive, live fish with large red patches and broad vertical or oblique red bands on body); the upper lobe of caudal fin with 4-6 red or brown bars (3 or 4 bars in subadults), while the lower caudal-fin lobe with 5-7 (3-5) red, brown or dark-brown bars, usually crossed, entirely or ony dorsally, by a yellow, beige, pale red or pale brown stripe; bars on both lobes of pupil width or less in adults, interrupted by pale, partly hyaline interspaces of similar width or slightly narrower; the lower caudal-lobe tip sometimes black; barbels are entirely yellow; with a single yellow, beige, or pale brown mid-lateral body stripe of pupil width from snout tip through eye to caudal-fin base; the stripe sometimes covered by 1-4 sections with 1-4 dark dots, the posteriormost group of dots behind second dorsal-fin base, often connecting to a dark saddle behind second dorsal fin; the first dorsal fin with 3-4 often fused, pale-red, pale-brown or beige stripes, the fin-tip area not darker pigmented while the second dorsal fin has 3-4 red or brown well-separated stripes with hyaline interspaces; pectoral fins hyaline, pelvic and anal fins weakly pigmented and partly hyaline; preserved fish is pale brown, often with remains of dark pigmentation deriving from caudal-fin bars, mid-lateral dots, saddle and/or second dorsal-fin distal stripe (Ref. 121834). Uiblein, F., D.C. Gledhill, D.A. Pavlov, T.A. Hoang and S. Shaheen 2019 Three new goatfishes of the genus <i>Upeneus</i> (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in <i>U. margarethae</i>. Zootaxa 4683(2):151-196. (Ref. 121834) FB-Morphology-69278 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 2021-01-11 00:00:00 2021-01-11 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=69278 Dorsal spines (total): 78; Dorsal soft rays (total): 9 Uiblein, F., D.C. Gledhill, D.A. Pavlov, T.A. Hoang and S. Shaheen 2019 Three new goatfishes of the genus <i>Upeneus</i> (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in <i>U. margarethae</i>. Zootaxa 4683(2):151-196. (Ref. 121834) FB-Distribution-69278 http://purl.org/dc/dcmitype/Text text/html Nicolas Bailly 2020-11-09 00:00:00 2025-11-05 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=69278 Western Pacific: S Indonesia to Singapore and Central Philippines; to the Gulf of Thailand, Vietnam, South China, Taiwan, and S Japan. Uiblein, F., D.C. Gledhill, D.A. Pavlov, T.A. Hoang and S. Shaheen 2019 Three new goatfishes of the genus <i>Upeneus</i> (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in <i>U. margarethae</i>. Zootaxa 4683(2):151-196. (Ref. 121834) FB-Size-69278 http://purl.org/dc/dcmitype/Text text/html Nicolas Bailly 2020-11-09 00:00:00 2021-01-22 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=69278 16.0 cm SL (male/unsexed; (Ref. 121834)) Uiblein, F., D.C. Gledhill, D.A. Pavlov, T.A. Hoang and S. Shaheen 2019 Three new goatfishes of the genus <i>Upeneus</i> (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in <i>U. margarethae</i>. Zootaxa 4683(2):151-196. (Ref. 121834) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=2 Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-69278 http://purl.org/dc/dcmitype/Text text/html Nicolas Bailly 2020-11-09 00:00:00 2021-01-22 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=69278 demersal; marine; depth range 6 - 24 m (Ref. 121834) Uiblein, F., D.C. Gledhill, D.A. Pavlov, T.A. Hoang and S. Shaheen 2019 Three new goatfishes of the genus <i>Upeneus</i> (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in <i>U. margarethae</i>. Zootaxa 4683(2):151-196. (Ref. 121834)