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	<dc:identifier>FB-7486</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7486</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Gobiiformes</dwc:Order>
	<dwc:Family>Oxudercidae</dwc:Family>
	<dwc:Genus>Periophthalmus</dwc:Genus>
	<dwc:ScientificName>Periophthalmus novemradiatus (Hamilton, 1822)</dwc:ScientificName>
	<commonName xml:lang='en'>Pearse's mudskipper</commonName>
	<commonName xml:lang='tl'>Bia</commonName>
	<commonName xml:lang='th'>Kachangtin</commonName>
	<commonName xml:lang='th'>กะจังตีน</commonName>
	<commonName xml:lang=''>九刺弹涂鱼</commonName>
	<commonName xml:lang=''>九刺彈塗魚</commonName>
	<synonym relationship='synonym'>Gobius novemradiatus Hamilton, 1822</synonym>
	<synonym relationship='synonym'>Periophthalmus novermradiatus (Hamilton, 1822)</synonym>
	<synonym relationship='synonym'>Periophthalmus pearsei Eggert, 1935</synonym>
	<synonym relationship='misapplied name'>Periophthalmus variabilis (non Eggert, 1935)</synonym>
	<synonym relationship='misapplied name'>Periophthalmus variabilis variabilis (non Eggert, 1935)</synonym>
	<dcterms:created>1993-02-23</dcterms:created>
	<dcterms:modified>2009-08-18</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=7486'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4833&amp;speccode=7486'>Talwar, P.K. and A.G. Jhingran0 Inland fishes of India and adjacent countries. Volume 2. A.A. Balkema, Rotterdam, i-xxii + 543-1158, 1 pl. (Ref. 4833)</reference>
	<reference isbn='0-7305-6374-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=5218&amp;speccode=7486'>Murdy, E.O.0 A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Rec. Aust. Mus., Suppl. 11:1-93. (Ref. 5218)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31184&amp;speccode=7486'>Martin, K.L.M. and C.R. Bridges0 Respiration in water and air. p. 54-78. In M.H. Horn, K.L.M. Martin and M.A. Chotkowski (eds.) Intertidal fishes. Life in two worlds. Academic Press. 399 p. (Ref. 31184)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31402&amp;speccode=7486'>Swedish Museum of Natural History0 Fish collection database of the Naturhistoriska riksmuseet (Swedish Museum of Natural History). Ichthyology Section, Department of Vertebrate Zoology, Swedish Museum of Natural History, Stockholm, Sweden. (Ref. 31402)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=7486'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=34530&amp;speccode=7486'>Rema Devi, K.0 Gobioids of Ennore estuary and its vicinity. Rec. zool. Surv. India 90(1-4):161-189. (Ref. 34530)</reference>
	<reference isbn='974-89954-6-1.' url='http://www.fishbase.org/references/FBRefSummary.php?id=37773&amp;speccode=7486'>Monkolprasit, S., S. Sontirat, S. Vimollohakarn and T. Songsirikul0 Checklist of Fishes in Thailand. Office of Environmental Policy and Planning, Bangkok, Thailand. 353 p. (Ref. 37773)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=7486'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40919&amp;speccode=7486'>Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45255&amp;speccode=7486'>Kapoor, D., R. Dayal and A.G. Ponniah0 Fish biodiversity of India. National Bureau of Fish Genetic Resources Lucknow, India.775 p. (Ref. 45255)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47438&amp;speccode=7486'>Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50614&amp;speccode=7486'>Dahanukar, N., R. Raut and A. Bhat0 Distribution, endemism and threat status of freshwater fishes in the Western Ghats of India. J. Biogeogr 31:123-136. (Ref. 50614)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=7486'>Riede, K.0 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference>
	<reference isbn='974-272-617-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=52823&amp;speccode=7486'>Fish Team of the Trang Project0 Illustrated fish fauna of a mangrove estuary at Sikao, southwestern Thailand. Trang Project for Biodiversity and Ecological Significance of Mangrove Estuaries in Southeast Asia, Rajamangala Institute of Technology and the University of Tokyo, Trang and Tokyo. 60p. (Ref. 52823)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=7486'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83599&amp;speccode=7486'>Jaafar, Z. and H.K. Larson0 A new species of mudskipper, &lt;i&gt;Periophthalmus takita&lt;/i&gt; (Teleostei: Gobiidae: Oxudercinae), from Australia, with a key to the genus. Zool. Sci. 25:946-952. (Ref. 83599)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83659&amp;speccode=7486'>Jaafar, Z., M. Perrig and L.M. Chou0 &lt;i&gt;Periophthalmus variabilis&lt;/i&gt; (Teleostei: Gobiidae: Oxudercinae), a valid species of mudskipper, and a re-diagnosis of &lt;i&gt;Periophthalmus novemradiatus&lt;/i&gt;. Zool. Sci. 26:309-314. (Ref. 83659)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87596&amp;speccode=7486'>Rajan, P.T., C.R. Sreeraj and T. Immanuel0 Fish fauna of coral reef, mangrove, freshwater, offshore and seagrass beds of Andaman and Nicobar Islands. Zoological Survey of India, Andaman and Nicobar Regional Centre, Haddo, Port Blair. (Ref. 87596)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126264&amp;speccode=7486'>Mahadevan, G. and V. Ravi0 Length‐weight relationship of two mudskippers (Gobiidae: Oxudercinae) &lt;i&gt;Boleophthalmus boddarti&lt;/i&gt; (Pallas, 1770) and &lt;i&gt;Periophthalmus novemradiatus&lt;/i&gt; (Hamilton, 1822) from Vellar estuary, Southeast India. J. Appl. Ichthyol. 34:1358-1360. (Ref. 126264)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1993-02-23 00:00:00</dcterms:created>
	<dcterms:modified>2009-08-18 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7486</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>An intertidal and amphibious air-breather that actively shuttles back and forth between rock pools and air (Ref. 31184).</dc:description>
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	<reference isbn='0-7305-6374-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=5218&amp;speccode=7486'>Murdy, E.O. 1989 A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Rec. Aust. Mus., Suppl. 11:1-93. (Ref. 5218)</reference></dataObject><dataObject>
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	<dcterms:created>2010-03-03 00:00:00</dcterms:created>
	<dcterms:modified>2010-03-03 00:00:00</dcterms:modified>
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	<dc:description>This is a sexually dimorphic species, with males possessing greatly elongate first spine of first dorsal fin, reaching eighth element of second dorsal fin when appressed; when preserved first dorsal fin brown or black with white distal edge, no spots; margin of first dorsal fin concave in males while concave to straight in females; second dorsal fin with two black bands, broader one located inframarginally and the other located medially, black spots basally on membrane between dorsal fin elements; pelvic fin with prominent frenum and inner rays fused for more than half of length. D1  modally IX; D2 modally I,12; A modally I,13; caudal fin rays modally 15 segmented rays, with first and last two rays unbranched; 62-67 lateral scale rows; 18-24 predorsal scales (Ref. 83659).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83659&amp;speccode=7486'>Jaafar, Z., M. Perrig and L.M. Chou 2009 &lt;i&gt;Periophthalmus variabilis&lt;/i&gt; (Teleostei: Gobiidae: Oxudercinae), a valid species of mudskipper, and a re-diagnosis of &lt;i&gt;Periophthalmus novemradiatus&lt;/i&gt;. Zool. Sci. 26:309-314. (Ref. 83659)</reference></dataObject><dataObject>
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	<dcterms:created>2010-03-03 00:00:00</dcterms:created>
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	<dc:description>Dorsal spines (total): 9 - 11; Dorsal soft rays (total): 112;  Analspines: 1; Analsoft rays: 12 - 13</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83659&amp;speccode=7486'>Jaafar, Z., M. Perrig and L.M. Chou 2009 &lt;i&gt;Periophthalmus variabilis&lt;/i&gt; (Teleostei: Gobiidae: Oxudercinae), a valid species of mudskipper, and a re-diagnosis of &lt;i&gt;Periophthalmus novemradiatus&lt;/i&gt;. Zool. Sci. 26:309-314. (Ref. 83659)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-7486</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1993-02-23 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7486</dc:source>
	<dc:description>Indian Ocean:  India.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83659&amp;speccode=7486'>Jaafar, Z., M. Perrig and L.M. Chou 2009 &lt;i&gt;Periophthalmus variabilis&lt;/i&gt; (Teleostei: Gobiidae: Oxudercinae), a valid species of mudskipper, and a re-diagnosis of &lt;i&gt;Periophthalmus novemradiatus&lt;/i&gt;. Zool. Sci. 26:309-314. (Ref. 83659)</reference></dataObject><dataObject>
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	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>fisheries: of no interest</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4833&amp;speccode=7486'>Talwar, P.K. and A.G. Jhingran 1991 Inland fishes of India and adjacent countries. Volume 2. A.A. Balkema, Rotterdam, i-xxii + 543-1158, 1 pl. (Ref. 4833)</reference></dataObject><dataObject>
	<dc:identifier>FB-Migration-7486</dc:identifier>
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	<dc:description>Amphidromous. Refers to fishes that regularly migrate between freshwater and the sea (in both directions), but not for the purpose of breeding, as in anadromous and catadromous species. Sub-division of diadromous.  Migrations should be cyclical and predictable and cover more than 100 km.Characteristic elements in amphidromy are: reproduction in fresh water, passage to sea by newly hatched larvae, a period of feeding and growing at sea usually a few months long, return to fresh water of well-grown juveniles, a further period of feeding and growing in fresh water, followed by reproduction there (Ref. 82692).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=7486'>Riede, K. 2004 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:description>10.0 cm SL (male/unsexed; (Ref. 4833))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4833&amp;speccode=7486'>Talwar, P.K. and A.G. Jhingran 1991 Inland fishes of India and adjacent countries. Volume 2. A.A. Balkema, Rotterdam, i-xxii + 543-1158, 1 pl. (Ref. 4833)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dcterms:created>1993-02-23 00:00:00</dcterms:created>
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	<dc:title>Environment</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7486</dc:source>
	<dc:description>demersal; amphidromous (Ref. 51243); brackish; marine</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=7486'>Riede, K. 2004 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference></dataObject>
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